diff --git a/projects/plugins/boost/app/features/image-guide/Image_Guide.php b/projects/plugins/boost/app/features/image-guide/Image_Guide.php
index 345f301c5f7bf..8d6d2ff5e9aa0 100644
--- a/projects/plugins/boost/app/features/image-guide/Image_Guide.php
+++ b/projects/plugins/boost/app/features/image-guide/Image_Guide.php
@@ -15,7 +15,7 @@ public function setup() {
 		}
 
 		// phpcs:ignore WordPress.Security.NonceVerification.Recommended
-		$override = defined( 'JETPACK_BOOST_IMAGE_GUIDE' ) && JETPACK_BOOST_IMAGE_GUIDE && isset( $_GET['jb-debug-ig'] );
+		$override = isset( $_GET['jb-debug-ig'] );
 
 		// Show the UI only when the user is logged in, with sufficient permissions and isn't looking at the dashboard.
 		if ( true !== $override && ( is_admin() || ! is_user_logged_in() || ! current_user_can( 'manage_options' ) ) ) {
diff --git a/projects/plugins/boost/app/features/optimizations/Optimizations.php b/projects/plugins/boost/app/features/optimizations/Optimizations.php
index 23ba293d38758..a39b39267f443 100644
--- a/projects/plugins/boost/app/features/optimizations/Optimizations.php
+++ b/projects/plugins/boost/app/features/optimizations/Optimizations.php
@@ -35,12 +35,9 @@ public function __construct() {
 			new $critical_css_class(),
 			new Lazy_Images(),
 			new Render_Blocking_JS(),
+			new Image_Guide(),
 		);
 
-		if ( defined( 'JETPACK_BOOST_IMAGE_GUIDE' ) && JETPACK_BOOST_IMAGE_GUIDE ) {
-			$features[] = new Image_Guide();
-		}
-
 		foreach ( $features as $feature ) {
 			$slug                    = $feature->get_slug();
 			$this->features[ $slug ] = new Optimization( $feature );
diff --git a/projects/plugins/boost/changelog/boost-enable-image-guide b/projects/plugins/boost/changelog/boost-enable-image-guide
new file mode 100644
index 0000000000000..0f6da559ae2c8
--- /dev/null
+++ b/projects/plugins/boost/changelog/boost-enable-image-guide
@@ -0,0 +1,4 @@
+Significance: patch
+Type: added
+
+Enable the Image Guide feature without a constant