Default Environment in project settings resets after project reloads and give errors

[ydeltastar]

Tested versions

• 4.4.dev
• Probably an effect of Universalize UID support in all resource types #97352 or Support uid:// in more places #99286

System information

Windows 11

Issue description

Project settings rendering/environment/defaults/default_environment resets when the project is reloaded.

After the project opens, the text field will be blank but the setting is still there in the project.godot pointing to an uid://.... The Environment is not applied in the 3D scene though. I get errors:

core\io\resource_uid.cpp:133 - Condition "!unique_ids.has(p_id)" is true. Returning: String()
Resource file not found: (expected type: )

Doesn't seem to happen with other path settings like gui/theme/custom.

Steps to reproduce

It only happens when the project has a lot of files so the editor takes a while on the first filesystem scan.

• Set a default Environment resource in Project Settings > rendering/environment/defaults/default_environment.
• Reload the project.
• Errors are printed, the environment is not applied in the scenes, and the field is blank in the settings.

Minimal reproduction project (MRP)

N/A